[1] Xia D P, Li J H, Garber P A, et al. Grooming reciprocity in male Tibetan macaques[J]. American journal of primatology, 2013, 75(10): 1009−1020. doi: 10.1002/ajp.22165
[2] 李保国,张鹏,渡边邦夫,等. 川金丝猴的相互理毛行为是否具有卫生功能[J]. 动物学报,2002,48(6):707−715.
[3]

Berthier J M, Semple S. Observing grooming promotes affiliation in Barbary macaques[J]. Proceedings of the Royal Society. B, Biological Sciences, 2018, 285(1893): 1−8.
[4]

Borgeaud C, Bshary R. Wild Vervet Monkeys Trade Tolerance and Specific Coalitionary Support for Grooming in Experimentally Induced Conflicts[J]. Current Biology, 2015, 25(22): 3011−3016. doi: 10.1016/j.cub.2015.10.016
[5]

Shutt K, Maclarnon A, Heistermann M, et al. Grooming in Barbary macaques: better to give than to receive?[J]. Biology Letters, 2007, 3(3): 231−233. doi: 10.1098/rsbl.2007.0052
[6]

Sonnweber R S, Massen J J M, Fitch W T. Post-copulatory grooming: a conditional mating strategy?[J]. Behavioral Ecology and Sociobiology, 2015, 69(11): 1749−1759. doi: 10.1007/s00265-015-1987-9
[7]

Kanngiesser P, Sueur C, Riedl K, et al. Grooming network cohesion and the role of individuals in a captive chimpanzee group[J]. American Journal of Primatology, 2011, 73(8): 758−767. doi: 10.1002/ajp.20914
[8]

Wei W, Qi X, Garber P A, et al. Supply and Demand Determine the Market Value of Access to Infants in the Golden Snub-Nosed Monkey (<italic>Rhinopithecus roxellana</italic>)[J]. PloS One, 2013, 8(6): e65962. doi: 10.1371/journal.pone.0065962
[9] 陈仕望,吴明阳,孙丙华,等. 黄山短尾猴理毛发生部位的差异[J]. 兽类学报,2021,41(3):330−337.
[10]

Ghiglieri M P. The Chimpanzees of Kibale Forest: a Field Study of Ecology and Social Structure[M]. New York: Columbia University Press, 1984.
[11]

Hamilton W D. The Genetical Evolution of Social Behavior[J]. Journal of Theoretical Biology, 1964, 7(1): 1−16. doi: 10.1016/0022-5193(64)90038-4
[12]

Noe R, Hammerstein P. Biological markets[J]. Trends in Ecology & Evolution, 1995, 10(8): 336−339.
[13]

Wei W, Qi X G, Guo S T, et al. Market Powers Predict Reciprocal Grooming in Golden Snub-Nosed Monkeys (<italic>Rhinopithecus roxellana</italic>)[J]. Plos One, 2012, 7(5): e36802. doi: 10.1371/journal.pone.0036802
[14]

Barrett L, Henzi S P. Monkeys, markets and minds: biological markets and primate sociality[M]. Cooperation in Primates and Human, 2006, 209-232.
[15] 李银华,李保国. 灵长类相互理毛的影响因素、功能及其利益分析[J]. 人类学学报,2004(4):334−342.
[16]

Goodall J. The Chimpanzees of Gombe: Patterns of Behavior[M]. Cambridge, Massachusetts: Belknap Press of Harvard University Press, 1986.
[17]

De Waal F B M, Luttrell L M. Mechanisms of Social Reciprocity in Three Primate Species: Symmetrical Relationship Characteristics or Cognition?[J]. Ethology & Sociobiology, 1988, 9(2-4): 101−108.
[18]

Schino G. Grooming, competition and social rank among female primates: a meta-analysis[J]. Animal Behaviour, 2001, 62(2): 265−271. doi: 10.1006/anbe.2001.1750
[19]

Franz C. Allogrooming behavior and grooming site preferences in captive bonobos (<italic>Pan paniscus</italic>): Association with female dominance[J]. International Journal of Primatology, 1999, 20(4): 525−546. doi: 10.1023/A:1020338706800
[20]

Long Y C, Kirkpatrick C R, Zhong T, et al. Report on the Distribution, Population, Ecology of the Yunnan Snub-nosed Monkey (<italic>Rhinopithecus bieti</italic>)[J]. Primates, 1994, 35(2): 241−250. doi: 10.1007/BF02382060
[21]

Li B G, Pan R L, Oxnard C E. Extinction of Snub-Nosed Monkeys in China During the Past 400 Years[J]. International Journal of Primatology, 2002, 23(6): 1227−1244. doi: 10.1023/A:1021122819845
[22]

Long Y C, Bleisch W V, Richardson M. Rhinopithecus bieti. The IUCN Red List of Threatened Species 2020: e. T19597A17943738. 2020.
[23]

Xiao W, Ding W, Cui L W, et al. Habitat Degradation ofRhinopithecus bieti in Yunnan, China[J]. International Journal of Primatology, 2003, 24(2): 389−398. doi: 10.1023/A:1023009518806
[24]

Huang Z P, Scott M B, Li Y P, et al. Black-and-white snub-nosed monkey (<italic>Rhinopithecus bieti</italic>) feeding behavior in a degraded forest fragment: clues to a stressed population[J]. Primates, 2017, 58(4): 517−524. doi: 10.1007/s10329-017-0618-7
[25]

Kirkpatrick R C, Long Y C, Zhong T, et al. Social organization and range use in the Yunnan snub-nosed monkey <italic>Rhinopithecus bieti</italic>[J]. International Journal of Primatology, 1998, 19(1): 13−51. doi: 10.1023/A:1020302809584
[26] 黎大勇,任宝平,和鑫明,等. 白马雪山自然保护区响古箐滇金丝猴的食性[J]. 兽类学报,2011,31(4):241−248.
[27]

Xiang Z F, Huo S, Xiao W. Activity budget of <italic>Rhinopithecus bieti</italic> at Tibet: Effects of day length, temperature and food availability[J]. Current Zoology, 2010, 56(6): 650−659. doi: 10.1093/czoolo/56.6.650
[28]

Wang S J, Huang Z P, He Y C, et al. Mating behavior and birth seasonality of black-and-white snub-nosed monkeys (<italic>Rhinopithecus bieti</italic>) at Mt. Lasha[J]. Zoological Research, 2012, 33(3): 241−248.
[29] 和鑫明,夏万才,巴桑,等. 滇金丝猴主雄应对配偶雌性数量的理毛策略[J]. 广西师范大学学报(自然科学版),2021,39(1):38−44.
[30]

Jeanne A. Observational study of behavior: Sampling methods[J]. Behaviour, 1974, 49(4): 227−265.
[31] 夏万才,胡杰,任宝平,等. 人工辅助投食滇金丝猴一雄多雌单元之间的等级序列[J]. 兽类学报,2017,4(37):57−64.
[32] 魏玮. 秦岭川金丝猴中互惠利他行为及生物市场现象的研究[D]. 西北大学, 2013.
[33]

Manson J H, David Navarrete C, Silk J B, et al. Time-matched grooming in female primates? New analyses from two species[J]. Animal Behaviour, 2004, 67(3): 493−500. doi: 10.1016/j.anbehav.2003.05.009
[34]

Cui L W, Sun Q L, Li B G. Dominance hierarchy and social relationships in a group of Captive black-and-white snub-nosed monkeys (<italic>Rhinopithecus bieti</italic>)[J]. Zoological Research, 2014, 35(3): 204−213.
[35]

Li Y P, Zhong T, Huang Z P, et al. Male and female birth attendance and assistance in a species of non-human primate (<italic>Rhinopithecus bieti</italic>)[J]. Behavioural Processes, 2020, 181: 104248. doi: 10.1016/j.beproc.2020.104248
[36]

Fairbanks L A. Relationships among adult females in captive vervet monkeys: Testing a model of rank-related attractiveness[J]. Animal Behaviour, 1980, 28(3): 853−859. doi: 10.1016/S0003-3472(80)80145-X